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Dissertation Title:  An Investigation into the Mechanism of Oscillatory Cyclic AMP Synthesis in Dictyostelium discoideum

Dissertation Title: An X-Ray Diffraction Study of Chloroplast Thylakoid Membrane Structure

PhD Dissertation Advisor:  Dr. Donald L.D. Caspar, Children's Cancer Research Foundation at Boston Children's Hospital

Dissertation Title: The Reactive Sulphydryl Groups of the Band 3 Polypeptide of Human Erythrocyte Membranes

PhD Dissertation Advisor: Dr. Guido Guidotti, Dept. of Biochemistry & Molecular Biology, Harvard University

Dissertation Title:  Rate Processes in the Terminal Events of Complement Hemolysis

PhD Dissertation Advisor: Dr. Donald L.D. Caspar, Children's Cancer Research Foundation at Boston Children's Hospital

Dissertation Title: The Primary Structure of the Calcium-dependent Adenosine Tri-phosphatase from Rabbit Muscle Sarcoplasmic Reticulum

PhD Dissertation Advisor: Dr. Guido Guidotti, Dept. of Biochemistry & Molecular Biology, Harvard University